(* indicates equal contribution, # indicates undergraduate author)



EM Caves and LA Kelley. 2023. Proportional processing of a visual mate choice signal in the green swordtail Xiphophorus helleriiEcology Letters 26: 575-585


EM Caves, PA Green, and M Hughes. 2022. Communication. In: The Routledge International Handbook of Comparative Psychology. Pg 147-159. Edited by: A.R. Ridley, P. d’Ettorre, and T.M. Freeberg. Routledge.


EM Caves, F de Busserolles, and LA Kelley. Sex differences in behavioural and anatomical estimates of visual acuity in the green swordtail Xiphophorus helleri. 2021. Journal of Experimental Biology 224: jeb243420.

T Dixit, EM Caves, CN Spottiswoode, and NPC Horrocks. Why and how to apply Weber’s Law to the coevolution of models and mimics. Evolution 75: 1906-1919. doi:10.1111/evo.14290

EM Caves. The behavioural ecology of marine cleaning mutualisms. 2021. Biological Reviews. doi: 10.1111/brv.12770

EM Caves,T Dixit, JFR Colebrook-Robjent, L Hamusikili, M Stevens, R Thorogood, and CN Spottiswoode. Hosts elevate either within-clutch consistency or between-clutch distinctiveness of egg phenotypes in defence against brood parasites. 2021. Proceedings of the Royal Society B: 20210326.

EM Caves and S Johnsen. 2021. The sensory impacts of climate change: bathymetric shifts and visually-mediated interactions in aquatic species. Proceedings of the Royal Society B. 288: 20210396. Covered by:, Energy & Environment News

EM Caves, PA Green, MN Zipple, D Bharath#, S Peters, S Johnsen, and S Nowicki. 2021. Comparison of categorical color perception in two Estrildid finches. The American Naturalist: 197: 190-202.


EM Caves, J Troscianko*, and LA Kelley*. 2020. A customizable, low-cost optomotor apparatus: a powerful tool for behaviourally measuring visual capability. Methods in Ecology and Evolution.

EM Caves, LE Schweikert, PA Green, C Taboada, MN Zipple, S Peters, S Nowicki, and S Johnsen. 2020. Variation in retinal carotenoids correlates with variation in perception of carotenoid coloration. Behavioral Ecology and Sociobiology. DOI:10.1007/s00265-020-02874-5


EM Caves*, C Chen*#, and S Johnsen. 2019. The cleaner shrimp Lysmata amboinensis adjusts its behavior toward predatory versus non-predatory clients. Biology Letters: 15, 20190534.

EM Caves, S Nowicki, and S Johnsen. 2019. Von Uexküll revisited: Addressing human biases in the study of animal perception. Integrative and Comparative Biology. DOI:10.1093/icb/icz073

MN Zipple*, EM Caves*, PA Green, S Peters, S Johnsen, and S Nowicki. 2019. Categorical perception in non-signalling versus signalling color ranges in a songbird. Proceedings of the Royal Society B: 286, 20190524

LE Schweikert, EM Caves, SE Solie, TT Sutton, and S Johnsen. 2019. Variation in rod spectral sensitivity of fishes is best predicted by habitat and depth. Journal of Fish Biology. 95: 179-185. DOI:10.1111/jfb.13859


LE Schweikert, RR Fitak, EM Caves, TT Sutton, and S Johnsen. 2018. Spectral sensitivity in ray-finned fishes: diversity, ecology, and shared descent. Journal of Experimental Biology: 221, jeb189761. doi:10.1242/jeb.189761

EM Caves*, PA Green*, MN Zipple, S Peters, S Johnsen, and S Nowicki. 2018. Categorical perception of colour signals in a songbird. Nature. 560: 365–367.

EM Caves, PA Green, and S Johnsen. 2018. Mutual visual signalling between the cleaner shrimp Ancylomenes pedersoni and its client fish. Proceedings of the Royal Society B 285: 20180800.

EM Caves, NC Brandley, and S Johnsen. 2018. Visual acuity and the evolution of signals. Trends in Ecology and Evolution. 33: 358-372.

R Fitak, EM Caves, and S Johnsen. 2018. Orientation in pill bugs: an interdisciplinary activity to engage students in concepts of biology, physics, and circular statistics. The American Biology Teacher 80: 608-618.


EM Caves and S Johnsen. 2017. AcuityView: An R package for portraying the effects of visual acuity on scenes observed by an animal. Methods in Ecology and Evolution 9: 793-797.

EM Caves, M Stevens, and CN Spottiswoode. 2017. Does coevolution with a shared parasite drive hosts to partition their defences among species? Proceedings of the Royal Society B. 284: 20170272.

EM Caves, TT Sutton, and S Johnsen. 2017. Visual acuity in fish correlates with eye size and habitat. Journal of Experimental Biology 220: 1586-1596.


EM Caves, TM Frank, and S Johnsen. 2016. Spectral sensitivity, spatial resolution, and temporal resolution and their implications for conspecific signalling in cleaner shrimp. Journal of Experimental Biology 219: 597-608.


EM Caves, M Stevens, E Iverson, and CN Spottiswoode. 2015. Hosts of brood parasites have evolved egg visual signatures with elevated information content. Proceedings of the Royal Society B 282: 20150598. *cover


EM Caves*, Jennings SB*, HilleRisLambers J, Tewksbury JJ, Rogers HS. 2013. Natural experiment demonstrates that bird loss leads to cessation of dispersal of native seeds from intact to degraded forests. PLoS ONE 8(5): e65618.